Diluvicursor

Diluvicursor ("flood runner") is a genus of small ornithischian from the Lower Albian (Early Cretaceous) of Australia. It is known from one species, the type species D. pickeringi. The two known specimens, a vertebra and a partial juvenile postcranium discovered in 2005 from the Eumeralla Formation, are known, and they were named in early 2018.[1]

Diluvicursor
Temporal range: Albian
~113–112 Ma
↓
Skeletal diagram showing the remains of Diluvicursor's holotype
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Order: †Ornithischia
Suborder: †Ornithopoda
Genus: †Diluvicursor
Herne et al., 2018
Type species
†Diluvicursor pickeringi
Herne et al., 2018

Discovery and naming

Map showing the location of the Eumeralla Formation, EF in C, and localities within the formation, B

The remains of Diluvicursor were discovered in 2005 in the Eumeralla Formation, dating from the early Albian, in Victoria, southeastern Australia at the beach of Cape Otway near the extensively excavated Dinosaur Cove, a location where the remains of several other ornithischians had been previously discovered, alongside remains of other species. The then new locality was named the "Eric the Red West Sandstone" (ETRW). The find was reported in the scientific literature by Thomas Rich et al., in 2009, in a study about remains from ETRW referred to the mammal Bishops. The authors tentatively it referred to Ornithopoda. The vertebra of a spinosaurid, NMV P221081, was also found at the site. In a 2013 review of ornithopods from Victoria, it was given a diagnosis and preliminary description.[1][2][3]

The type specimen, preserving a partial tail and hindlimb

The specimens were described and named online, through use of ZooBank registration, in January 2018, in a study in which the sedimentology, taphonomy, palaeoecology, and stratigraphy of the site were also investigated. The generic name of the taxon is derived from the Latin diluvi~, meaning "flood" or "deluge", which was chosen in reference to the fact that the type specimen was deposited in a high energy river and the species would have lived in a prehistoric floodplain, and the Latin word cursor, which means "runner". The specific name, pickeringi, is honoring David A. Pickering, who had made notable contributions to palaeontology in Australia, and who died at the time of the preparation of the study.[1][4]

The holotype, NMV P221080, consists of an almost complete articulated tail lacking only the first, second and twelfth tail vertebra, and parts of the lower right leg: the lower end of the shinbone and calfbone, the complete ankle bones, the metatarsus, the first toe and the first phalanges of the second, third and fourth toes. This specimen is preserved on a stone plate, separated into five blocks. It would have been deposited in a deep and high-energy river, the carcass thought to have been caught in a trap of plant debris on a standing tree stump. It represents a juvenile individual. Specimen NMV P229456, a partial tail vertebra of a larger individual, found nearby in the same deposit, was referred to the taxon.[1]

Description

Life restoration of Diluvicursor in its environment

The holotype individual, thought to be an older juvenile, was estimated to be 1.2 metres (3.9 ft) in length, using the proportions of Hypsilophodon to fill in unknown areas; it has been compared to the size of a modern wild turkey. It was noted, however, that the tail being preserved in a curled fashion made exact tail length uncertain, owing to difficulty in measuring intervertebral space; the size estimate is therefore uncertain. Based on an isolated caudal vertebra, NMV P229456, twice the length of vertebrae from the holotype, adults are estimated to have measured at least 2.3 metres (7.5 ft) long, closer to the size of a rhea, and still small for an ornithopod.[1] The taxon was noted in Herne's PhD thesis as being distinctly more robust than other assumed relatives from the area.[3] Ten autapomorphies, unique traits, were identified in the holotype.[1]

Phylogeny

The cladogram below results from analysis by Herne et al., 2019.[5]

Ornithischia
 

Heterodontosauridae

 
 
 

Eocursor

 
 
 

Thyreophora

 
Neornithischia
 

Lesothosaurus

 
 
 

Agilisaurus

 
 
 
 

Hexinlusaurus

 
 

Yandusaurus

 
 
 
 
 

Nanosaurus

 
 
Jeholosauridae
 

Haya

 
 
 

Jeholosaurus

 
 

Changchunsaurus

 
 
 
Thescelosauridae
 
 

Orodromeus

 
 

Koreanosaurus

 
 
 
 

Zephyrosaurus

 
 
 

Yueosaurus

 
 

Thescelosaurus

 
 
 
 
 
 
Cerapoda
 

Marginocephalia

 
Ornithopoda
 

Parksosaurus

 
 
Elasmaria
 

Talenkauen

 
 
 
 

Macrogryphosaurus

 
 

Gasparinisaura

 
 
 
 

Galleonosaurus

 
 

Leaellynasaura

 
 

Anabisetia

 
 

Diluvicursor

 
 
 
 
Clypeodonta
 

Hypsilophodon

 
Iguanodontia
 

Rhabdodontidae

 
 
 

Muttaburrasaurus

 
 
 

Tenontosaurus

 
 

Dryomorpha

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 

See also

References

  1. Herne, M.C.; Tait, A.M; Weisbecker, V.; Hall, M.; Nair, J.P; Cleeland, M.; Salisbury, S.W. (2018). "A new small-bodied ornithopod (Dinosauria, Ornithischia) from a deep, high-energy Early Cretaceous river of the Australian–Antarctic rift system". PeerJ. 5: e4113. doi:10.7717/peerj.4113. PMC 5767335. PMID 29340228.
  2. Rich T.H., Vickers-Rich P., Flannery T.F., Pickering D., Kool L., Tait A.M., Fitzgerald E.M.G. 2009. "A fourth Australian Mesozoic mammal locality". Museum of Northern Arizona, Bulletin 65: 677–681
  3. Herne, M. (2013). "Anatomy, systematics and phylogenetic relationships of the Early Cretaceous ornithopod dinosaurs of the Australian-Antarctic rift system" PhD Thesis. The University of Queensland
  4. Kool, Lesley. "Farewell David Pickering". Dinosaur Dreaming. Retrieved June 20, 2018.
  5. Herne, Matthew C.; Nair, Jay P.; Evans, Alistair R.; Tait, Alan M. (2019). "New small-bodied ornithopods (Dinosauria, Neornithischia) from the Early Cretaceous Wonthaggi Formation (Strzelecki Group) of the Australian-Antarctic rift system, with revision of Qantassaurus intrepidus Rich and Vickers-Rich, 1999". Journal of Paleontology. doi:10.1017/jpa.2018.95.
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